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Arthur Berg, PhD

Arthur Berg, PhD

Professor, Department of Public Health Sciences
Division of Biostatistics and Bioinformatics
Professor, Department of Surgery
Professor, Department of Neurosurgery
Professor, Department of Family and Community Medicine
Professor, Statistics
Associate Professor, Department of Public Health Sciences
Associate Professor, Department of Surgery
Scientific Program:Cancer Control

Research Interests

Dr. Arthur Berg frequently engages in collaborative research relating to all forms of biostatistics and bioinformatics analyses. He has utilized numerous software tools to perform diverse types of statistical analyses, and he has developed many computational tools of his own.

In addition to his computational strengths, Dr. Berg has also written a number of methodological papers in biostatistics, statistics and statistical genetics.

During his time at Penn State, Dr. Berg has participated in several successful collaborations with faculty, postdocs and students that has involved biostatistical expertise.

  • Genes
  • Neoplasms
  • Single Nucleotide Polymorphism
  • Crohn Disease
  • Population
  • Spinal Cord Injuries
  • Inflammatory Bowel Diseases
  • Gene Expression
  • Mutation
  • Ulcerative Colitis
  • Alleles
  • Haplotypes

Recent Publications


Farkas, GJ, Cunningham, PM, Sneij, AM, Hayes, JE, Nash, MS, Berg, AS, Gater, DR & Rolls, BJ 2024, 'Reasons for meal termination, eating frequency, and typical meal context differ between persons with and without a spinal cord injury', Appetite, vol. 192, 107110.
Onks, C, Weaver, L, Latorre, J, Silvis, M, Berg, A, Phillips, S, Loeffert, J, French, C & Armstrong, A 2024, 'The most effective corticosteroid dose in the treatment of glenohumeral osteoarthritis: Feasibility pilot and protocol for double blinded randomized controlled trial', Osteoarthritis and Cartilage Open, vol. 6, no. 3, 100484.


Darby, A, Parascando, JA, Lipinski, M, Lipinski, C, Mendez-Miller, M, Berg, A, Rabago, D & Oser, TK 2023, 'Awareness and interest in osteopathic manipulative treatment in allopathic medical students', Journal of Osteopathic Medicine, vol. 123, no. 8, pp. 379-384.
Tom, CM, Mankarious, MM, Jeganathan, NA, Deutsch, M, Koltun, WA, Berg, AS & Scow, JS 2023, 'Characteristics and Outcomes of Right- Versus Left-Sided Early-Onset Colorectal Cancer', Diseases of the colon and rectum, vol. 66, no. 4, pp. 498-510.
Karpa, K, Ward, J, Stegman, M, Berg, A & Leong, SL 2023, 'IPEx: A gamification tool for learner application of pharmacologic principles of opioid use, misuse, and addiction', Pharmacology Research and Perspectives, vol. 11, no. 5, e01141.
Larsen, M, He, F, Kawasawa, YI, Berg, A, Vgontzas, AN, Liao, D, Bixler, EO & Fernandez-Mendoza, J 2023, 'Objective and subjective measures of sleep initiation are differentially associated with DNA methylation in adolescents', Clinical Epigenetics, vol. 15, no. 1, 136.
Mankarious, MM, Hughes, AJ, Berg, AS, Scow, JS, Jeganathan, AN, Kulaylat, AS & Deutsch, MJ 2023, 'Survival outcomes of anal adenocarcinoma versus rectal adenocarcinoma: A retrospective cohort study', Indian Journal of Gastroenterology, vol. 42, no. 5, pp. 694-700.
Walser, SA, Costigan, H, Stuckey, HL, Berg, A & Stephens, MB 2023, 'The Opaque Language of Sexuality: Medical Students’ and Providers’ Beliefs About Virginity', Archives of Sexual Behavior, vol. 52, no. 6, pp. 2619-2638.
Farkas, GJ, Berg, AS, Sneij, A, Dolbow, DR & Gorgey, AS 2024, 'The comparison of total energy and protein intake relative to estimated requirements in chronic spinal cord injury', British Journal of Nutrition, vol. 131, no. 3, pp. 489-499.
Wang, H, Berger, KN, Miller, EL, Fu, W, Broglie, L, Goldman, FD, Konig, H, Lim, SJ, Berg, AS, Talano, JA, Comito, MA, Farag, SS & Pu, JJ 2023, 'The impacts of total body irradiation on umbilical cord blood hematopoietic stem cell transplantation', Therapeutic Advances in Hematology, vol. 14.
Satriale, F, Garman, JC, Roy, S, Parascando, J, Berg, A & Oser, T 2023, 'Understanding Nutritional Knowledge and Experiences in Families With a Child Newly Diagnosed With Type 1 Diabetes', Diabetes Spectrum, vol. 36, no. 1, pp. 52-58.


Riley, TD, Radico, JA, Parascando, J, Berg, A & Oser, TK 2022, 'Challenges in Effective Faculty and Provider Recognition to Enhance Engagement', Family medicine, vol. 54, no. 6, pp. 461-465.
Schlegel, D, Heron, C, Berg, A, Parascando, J & Silvis, M 2022, 'Diagnoses per Encounter by Telephone, Televideo, and In-Office Visits', Journal of the American Board of Family Medicine, vol. 35, no. 3, pp. 491-496.
Hawila, N & Berg, A 2023, 'Exact-corrected confidence interval for risk difference in noninferiority binomial trials', Biometrics, vol. 79, no. 2, pp. 1133-1144.
Dolbow, DR, Farkas, GJ, Berg, AS, Welsch, MA, Gorgey, AS & Gater, DR 2022, 'Fat to lean mass ratio in spinal cord injury: Possible interplay of components of body composition that may instigate systemic inflammation and metabolic syndrome', Journal of Spinal Cord Medicine, vol. 45, no. 6, pp. 833-839.
Dinavahi, SS, Chen, YC, Gowda, R, Dhanyamraju, PK, Punnath, K, Desai, D, Berg, A, Kimball, SR, Amin, S, Yang, JM & Robertson, GP 2022, 'Targeting Protein Translation in Melanoma by Inhibiting EEF-2 Kinase Regulates Cholesterol Metabolism though SREBP2 to Inhibit Tumour Development', International journal of molecular sciences, vol. 23, no. 7, 3481.
Dinavahi, SS, Chen, YC, Punnath, K, Berg, A, Herlyn, M, Foroutan, M, Huntington, ND & Robertson, GP 2022, 'Targeting WEE1/AKT Restores p53-Dependent Natural Killer–Cell Activation to Induce Immune Checkpoint Blockade Responses in “Cold” Melanoma', Cancer Immunology Research, vol. 10, no. 6, pp. 757-769.


Chen, YC, Dinavahi, SS, Feng, Q, Gowda, R, Ramisetti, S, Xia, X, LaPenna, KB, Chirasani, VR, Cho, SH, Hafenstein, SL, Battu, MB, Berg, A, Sharma, AK, Kirchhausen, T, Dokholyan, NV, Amin, S, He, P & Robertson, GP 2021, 'Activating Sphingosine-1-phospahte signaling in endothelial cells increases myosin light chain phosphorylation to decrease endothelial permeability thereby inhibiting cancer metastasis', Cancer Letters, vol. 506, pp. 107-119.
Farkas, GJ, Swartz, AM, Gorgey, AS, Berg, AS & Gater, DR 2022, 'Acute exercise improves glucose effectiveness but not insulin sensitivity in paraplegia', Disability and Rehabilitation, vol. 44, no. 17, pp. 4656-4662.
Hawila, N & Berg, A 2021, 'Assessing the impact of COVID-19 on registered interventional clinical trials', Clinical and Translational Science, vol. 14, no. 3, pp. 1147-1154.
Berg, A 2021, 'Bayesian explorations with dice', Teaching Statistics, vol. 43, no. 3, pp. 114-123.
Gater, DR, Farkas, GJ, Dolbow, DR, Berg, A & Gorgey, AS 2021, 'Body Composition and Metabolic Assessment after Motor Complete Spinal Cord Injury: Development of a Clinically Relevant Equation to Estimate Body Fat', Topics in Spinal Cord Injury Rehabilitation, vol. 27, no. 1, pp. 11-22.
Kline, BP, Yochum, GS, Brinton, DL, Schieffer, KM, Weaver, T, Harris, L, Deiling, S, Berg, AS & Koltun, WA 2021, 'COLQ and ARHGAP15 are associated with diverticular disease and are expressed in the colon', Journal of Surgical Research, vol. 267, pp. 397-403.
Lafferty, L, Wawrzyniak, J, Chambers, M, Pagliarulo, T, Berg, A, Hawila, N & Silvis, M 2022, 'Clinical Indoor Running Gait Analysis May Not Approximate Outdoor Running Gait Based on Novel Drone Technology', Sports Health, vol. 14, no. 5, pp. 710-716.
Kline, BP, Weaver, T, Brinton, DL, Harris, L, Yochum, GS, Berg, AS & Koltun, WA 2021, 'Clinical and Genetic Factors Impact Time to Surgical Recurrence After Ileocolectomy for Crohn's Disease', Annals of surgery, vol. 274, no. 2, pp. 346-351.
Farkas, GJ, Gordon, PS, Trewick, N, Gorgey, AS, Dolbow, DR, Tiozzo, E, Berg, AS & Gater, DR 2021, 'Comparison of various indices in identifying insulin resistance and diabetes in chronic spinal cord injury', Journal of Clinical Medicine, vol. 10, no. 23, 5591.
Berg, A 2021, 'Concursos de modelos bayesianos para el salón de clases', Revista Colombiana de Estadistica, vol. 44, no. 2, pp. 243-252.
Golla, U, Ehudin, MA, Annageldiyev, C, Zeng, Z, Bastihalli Tukaramrao, D, Tarren, A, Date, AA, Elcheva, I, Berg, A, Amin, S, Loughran, TP, Kester, M, Desai, D, Dovat, S, Claxton, D & Sharma, A 2021, 'Dj4 targets the rho-associated protein kinase pathway and attenuates disease progression in preclinical murine models of acute myeloid leukemia', Cancers, vol. 13, no. 19, 4889.
Farkas, GJ, Gorgey, AS, Dolbow, DR, Berg, AS & Gater, DR 2021, 'Energy Expenditure, Cardiorespiratory Fitness, and Body Composition Following Arm Cycling or Functional Electrical Stimulation Exercises in Spinal Cord Injury: A 16-Week Randomized Controlled Trial', Topics in Spinal Cord Injury Rehabilitation, vol. 27, no. 1, pp. 121-134.
Batra, EK, Palusci, VJ & Berg, A 2022, 'Factors Associated with Child Maltreatment Fatality among Young Children with an Open Child Protective Services Case at Death', Child Abuse Review, vol. 31, no. 2, CAR2734.
Radico, J, Oser, T, Fausnight, T, Berg, A, Ouyang, A & Leong, SL 2021, 'Factors that Influence Work Family Conflict for Women Faculty [version 1]', MedEd Publish.
Veldheer, S, Scartozzi, C, Bordner, CR, Opara, C, Williams, B, Weaver, L, Rodriguez, D, Berg, A & Sciamanna, C 2021, 'Impact of a Prescription Produce Program on Diabetes and Cardiovascular Risk Outcomes', Journal of Nutrition Education and Behavior, vol. 53, no. 12, pp. 1008-1017.
Raca, G, Abdel-Azim, H, Yue, F, Broach, J, Payne, JL, Reeves, ME, Gowda, C, Schramm, J, Desai, D, Dovat, E, Hu, T, Berg, AS, Bhojwani, D, Payne, KJ & Dovat, S 2021, 'Increased Incidence of IKZF1 deletions and IGH-CRLF2 translocations in B-ALL of Hispanic/Latino children—a novel health disparity', Leukemia, vol. 35, no. 8, pp. 2399-2402.
GWAS and Sequencing Consortium of Alcohol and Nicotine Use (GSCAN) 2021, 'Model-based assessment of replicability for genome-wide association meta-analysis', Nature communications, vol. 12, no. 1, 1964.
Partin, M, Sanchez, A, Poulson, J, Berg, A, Parascando, J & Ramirez, SI 2021, 'Social inequities between prenatal patients in family medicine and obstetrics and gynecology with similar outcomes', Journal of the American Board of Family Medicine, vol. 34, no. 1, pp. 181-188.
Berg, A & Hawila, N 2021, 'Some teaching resources using R with illustrative examples exploring COVID-19 data', Teaching Statistics, vol. 43, no. S1, pp. S98-S109.


Liao, JG, Berg, A & McMurry, TL 2021, 'A Robustified Posterior for Bayesian Inference on a Large Number of Parallel Effects', American Statistician, vol. 75, no. 2, pp. 145-151.
Berg, A 2020, 'Assessing the Impact of the NIH CTSA Program on Clinical Trials Registered With', Clinical and Translational Science, vol. 13, no. 4, pp. 818-825.
Berg, A 2020, 'Bayesian statistics in the classroom: Introducing shrinkage with basketball statistics and the internet movie database', Teaching Statistics, vol. 42, no. 2, pp. 47-53.
Kline, BP, Weaver, T, Brinton, DL, Deiling, S, Yochum, GS, Berg, AS & Koltun, WA 2020, 'Clinical and Genetic Factors Associated With Complications After Crohn's Ileocolectomy', Diseases of the colon and rectum, vol. 63, no. 3, pp. 357-364.
Liao, JG, Midya, V & Berg, A 2021, 'Connecting and Contrasting the Bayes Factor and a Modified ROPE Procedure for Testing Interval Null Hypotheses', American Statistician, vol. 75, no. 3, pp. 256-264.
Dinavahi, SS, Gowda, R, Gowda, K, Bazewicz, CG, Chirasani, VR, Battu, MB, Berg, A, Dokholyan, NV, Amin, S & Robertson, GP 2020, 'Development of a Novel Multi-Isoform ALDH Inhibitor Effective as an Antimelanoma Agent', Molecular cancer therapeutics, vol. 19, no. 2, pp. 447-459.
Noel, OF, Berg, A, Onyango, N & MacKay, DR 2020, 'Ethnic and Gender Diversity Comparison between Surgical Patients and Caring Surgeons', Plastic and Reconstructive Surgery - Global Open, vol. 8, no. 10, pp. E3198.
Riley, TD, Curry, WJ, Beck, MJ, Berg, A & Ruffin, MT 2020, 'Feasibility of nonanonymous burnout surveys in a large academic department', Family medicine, vol. 54, no. 4, pp. 278-281.
Farkas, GJ, Gordon, PS, Swartz, AM, Berg, AS & Gater, DR 2020, 'Influence of mid and low paraplegia on cardiorespiratory fitness and energy expenditure', Spinal cord series and cases, vol. 6, no. 1, 110.
Liu, R, Onks, C, Petscavage-Thomas, J, Walker, E, Berg, A, Silvis, M & Dhawan, A 2021, 'Prevalence of shoulder labral abnormalities on MRI in a non-athletic asymptomatic young adult population', Skeletal Radiology, vol. 50, no. 5, pp. 921-925.
Berg, A 2021, 'Statistical Analysis of the International Mathematical Olympiad', Mathematical Intelligencer, vol. 43, no. 1, pp. 105-112.