Researcher Profile

Researcher Profile

Arthur Berg, PhD

Arthur Berg, PhD

Associate Professor, Department of Public Health Sciences
Division of Biostatistics and Bioinformatics
Associate Professor, Department of Surgery
Center for Statistical Genetics
Scientific Program:Cancer Control

Research Interests

Dr. Arthur Berg frequently engages in collaborative research relating to all forms of biostatistics and bioinformatics analyses. He has utilized numerous software tools to perform diverse types of statistical analyses, and he has developed many computational tools of his own.

In addition to his computational strengths, Dr. Berg has also written a number of methodological papers in biostatistics, statistics and statistical genetics.

During his time at Penn State, Dr. Berg has participated in several successful collaborations with faculty, postdocs and students that has involved biostatistical expertise.

  • Genes
  • Single Nucleotide Polymorphism
  • Neoplasms
  • Crohn Disease
  • Population
  • Spinal Cord Injuries
  • Inflammatory Bowel Diseases
  • Gene Expression
  • Mutation
  • Ulcerative Colitis
  • Alleles
  • Haplotypes

Recent Publications


Chen, YC, Dinavahi, SS, Feng, Q, Gowda, R, Ramisetti, S, Xia, X, LaPenna, KB, Chirasani, VR, Cho, SH, Hafenstein, SL, Battu, MB, Berg, A, Sharma, AK, Kirchhausen, T, Dokholyan, NV, Amin, S, He, P & Robertson, GP 2021, 'Activating Sphingosine-1-phospahte signaling in endothelial cells increases myosin light chain phosphorylation to decrease endothelial permeability thereby inhibiting cancer metastasis', Cancer Letters, vol. 506, pp. 107-119.
Farkas, GJ, Swartz, AM, Gorgey, AS, Berg, AS & Gater, D 2021, 'Acute exercise improves glucose effectiveness but not insulin sensitivity in paraplegia', Disability and Rehabilitation.
Hawila, N & Berg, A 2021, 'Assessing the impact of COVID-19 on registered interventional clinical trials', Clinical and Translational Science, vol. 14, no. 3, pp. 1147-1154.
Berg, A 2021, 'Bayesian explorations with dice', Teaching Statistics, vol. 43, no. 3, pp. 114-123.
Gater, DR, Farkas, GJ, Dolbow, DR, Berg, A & Gorgey, AS 2021, 'Body Composition and Metabolic Assessment After Motor Complete Spinal Cord Injury: Development of a Clinically Relevant Equation to Estimate Body Fat', Topics in Spinal Cord Injury Rehabilitation, vol. 27, no. 1, pp. 11-22.
Kline, BP, Yochum, GS, Brinton, DL, Schieffer, KM, Weaver, T, Harris, L, Deiling, S, Berg, AS & Koltun, WA 2021, 'COLQ and ARHGAP15 are associated with diverticular disease and are expressed in the colon', Journal of Surgical Research, vol. 267, pp. 397-403.
Farkas, GJ, Gorgey, AS, Dolbow, DR, Berg, AS & Gater, DR 2021, 'Energy Expenditure, Cardiorespiratory Fitness, and Body Composition Following Arm Cycling or Functional Electrical Stimulation Exercises in Spinal Cord Injury: A 16-Week Randomized Controlled Trial', Topics in Spinal Cord Injury Rehabilitation, vol. 27, no. 1, pp. 121-134.
Raca, G, Abdel-Azim, H, Yue, F, Broach, J, Payne, JL, Reeves, ME, Gowda, C, Schramm, J, Desai, D, Dovat, E, Hu, T, Berg, AS, Bhojwani, D, Payne, KJ & Dovat, S 2021, 'Increased Incidence of IKZF1 deletions and IGH-CRLF2 translocations in B-ALL of Hispanic/Latino children—a novel health disparity', Leukemia, vol. 35, no. 8, pp. 2399-2402.
GWAS and Sequencing Consortium of Alcohol and Nicotine Use (GSCAN) 2021, 'Model-based assessment of replicability for genome-wide association meta-analysis', Nature communications, vol. 12, no. 1, 1964.
Partin, M, Sanchez, A, Poulson, J, Berg, A, Parascando, J & Ramirez, SI 2021, 'Social inequities between prenatal patients in family medicine and obstetrics and gynecology with similar outcomes', Journal of the American Board of Family Medicine, vol. 34, no. 1, pp. 181-188.
Berg, A & Hawila, N 2021, 'Some teaching resources using R with illustrative examples exploring COVID-19 data', Teaching Statistics, vol. 43, no. S1, pp. S98-S109.


Liao, JG, Berg, A & McMurry, TL 2021, 'A Robustified Posterior for Bayesian Inference on a Large Number of Parallel Effects', American Statistician, vol. 75, no. 2, pp. 145-151.
Berg, A 2020, 'Assessing the Impact of the NIH CTSA Program on Clinical Trials Registered With', Clinical and Translational Science, vol. 13, no. 4, pp. 818-825.
Berg, A 2020, 'Bayesian statistics in the classroom: Introducing shrinkage with basketball statistics and the internet movie database', Teaching Statistics, vol. 42, no. 2, pp. 47-53.
Kline, BP, Weaver, T, Brinton, DL, Deiling, S, Yochum, GS, Berg, AS & Koltun, WA 2020, 'Clinical and Genetic Factors Associated With Complications After Crohn's Ileocolectomy', Diseases of the colon and rectum, vol. 63, no. 3, pp. 357-364.
Liao, JG, Midya, V & Berg, A 2021, 'Connecting and Contrasting the Bayes Factor and a Modified ROPE Procedure for Testing Interval Null Hypotheses', American Statistician, vol. 75, no. 3, pp. 256-264.
Dinavahi, SS, Gowda, R, Gowda, K, Bazewicz, CG, Chirasani, VR, Battu, MB, Berg, A, Dokholyan, NV, Amin, S & Robertson, GP 2020, 'Development of a Novel Multi-Isoform ALDH Inhibitor Effective as an Antimelanoma Agent', Molecular cancer therapeutics, vol. 19, no. 2, pp. 447-459.
Noel, OF, Berg, A, Onyango, N & MacKay, DR 2020, 'Ethnic and Gender Diversity Comparison between Surgical Patients and Caring Surgeons', Plastic and Reconstructive Surgery - Global Open.
Riley, TD, Curry, WJ, Beck, MJ, Berg, A & Ruffin, MT 2020, 'Feasibility of nonanonymous burnout surveys in a large academic department', Family medicine, vol. 54, no. 4, pp. 278-281.
Farkas, GJ, Gordon, PS, Swartz, AM, Berg, AS & Gater, D 2020, 'Influence of mid and low paraplegia on cardiorespiratory fitness and energy expenditure', Spinal cord series and cases, vol. 6, no. 1, 110.
Liu, R, Onks, C, Petscavage-Thomas, J, Walker, E, Berg, A, Silvis, M & Dhawan, A 2021, 'Prevalence of shoulder labral abnormalities on MRI in a non-athletic asymptomatic young adult population', Skeletal Radiology, vol. 50, no. 5, pp. 921-925.
Berg, A 2021, 'Statistical Analysis of the International Mathematical Olympiad', Mathematical Intelligencer, vol. 43, no. 1, pp. 105-112.


He, F, Berg, A, Imamura Kawasawa, Y, Bixler, EO, Fernandez-Mendoza, J, Whitsel, EA & Liao, D 2019, 'Association between DNA methylation in obesity-related genes and body mass index percentile in adolescents', Scientific reports, vol. 9, no. 1, 2079.
Farkas, GJ, Gorgey, AS, Dolbow, DR, Berg, AS & Gater, DR 2019, 'Caloric Intake Relative to Total Daily Energy Expenditure Using a Spinal Cord Injury-Specific Correction Factor: An Analysis by Level of Injury', American journal of physical medicine & rehabilitation, vol. 98, no. 11, pp. 947-952.
Farkas, GJ, Pitot, MA, Berg, AS & Gater, DR 2019, 'Correction: Nutritional status in chronic spinal cord injury: a systematic review and meta-analysis (Spinal Cord, (2019), 57, 1, (3-17), 10.1038/s41393-018-0218-4)', Spinal Cord, vol. 57, no. 3, pp. 256.
Riley, T, Radico, J, Parascando, J, Berg, A & Oser, T 2019, 'Gaps in Recognition Among Family Medicine Providers: An Opportunity to Reduce Work-Home Conflict and Improve Retention', Paper presented at 47th NAPCRG Annual Meeting, 11/16/19 - 11/20/19.
Oser, SM, Stuckey, HL, Parascando, JA, McGinley, EL, Berg, A & Oser, TK 2019, 'Glycated hemoglobin differences among blog-reading adults with type 1 diabetes compared with those who do not read blogs: Cross-sectional study', JMIR Diabetes, vol. 4, no. 2, e13634.
Radico, J, Oser, T, Fausnight, T, Ouyang, A, Berg, A, Dong, H & Leong, SL 2019, 'Interplay between Culture, Work-to-Family Conflict, and Work Demands in Women in Primary Care', Paper presented at STFM Annual Spring Conference, 5/27/19 - 6/1/19.
Sheehan, JP, Grills, I, Chiang, VL, Dong, H, Berg, A, Warnick, RE, Kondziolka, D & Kavanagh, B 2019, 'Quality of life outcomes for brain metastasis patients treated with stereotactic radiosurgery: pre-procedural predictive factors from a prospective national registry', Journal of neurosurgery, vol. 131, no. 6, pp. 1848-1854.
Berg, A, Politis, D, Suaray, K & Zeng, H 2020, 'Reduced bias nonparametric lifetime density and hazard estimation', Test, vol. 29, no. 3, pp. 704-727.
Matthews, SM, Eshelman, MA, Berg, AS, Koltun, WA & Yochum, GS 2019, 'The Crohn’s disease associated SNP rs6651252 impacts MYC gene expression in human colonic epithelial cells', PloS one, vol. 14, no. 2, e0212850.
Brunke-Reese, D, Ssentongo, P, Ssentongo, AE, Phillips, BE, Pauli, EM, Berg, A, Kelleher, SL & Soybel, DI 2019, 'The Role of Genetic Variant rs13266634 in SLC30A8/ZnT8 in Postoperative Hyperglycemia after Major Abdominal Surgery', Journal of Clinical Endocrinology and Metabolism, vol. 104, no. 9, pp. 3877-3892.


Dinavahi, SS, Noory, MA, Gowda, R, Drabick, J, Berg, A, Neves, R & Robertson, G 2018, 'Moving synergistically acting drug combinations to the clinic by comparing sequential versus simultaneous drug administrations', Molecular pharmacology, vol. 93, no. 3, pp. 190-196.
Farkas, GJ, Pitot, MA, Berg, AS & Gater, DR 2019, 'Nutritional status in chronic spinal cord injury: a systematic review and meta-analysis', Spinal Cord, vol. 57, no. 1, pp. 3-17.
Yingst, JM, Veldheer, S, Hrabovsky, S, Hammett, E, Nicholson, J, Berg, A & Foulds, J 2018, 'Pilot Randomized Trial of an Automated Smoking Cessation Intervention via Mobile Phone Text Messages as an Adjunct to Varenicline in Primary Care', Journal of Health Communication, vol. 23, no. 4, pp. 370-378.
Rice, L, Sung, JH, Keane, K, Peterson, E, Sosnoff, J, Farkas, G, Swartz, A, Strath, S, Gorgey, A, Berg, A, Gater, D, Dyson-Hudson, T, Malanga, G, Cherian, C, Michalec, M, Kirshblum, S, Miller, C, Garlanger, K, Kortes, S, Schnorenberg, A, Slavens, B, Lee, K, Potter-Baker, K, Frost, F, Plow, E, Solinsky, R, Wilson, C, Henry, CA, Lombard, A, Maher, M, Weir, J, Saeed, S, Cirnigliaro, C, Specht, A, Garbarini, E, Augustine, J, Forrest, G, Bauman, W, Maher, M, Lombard, A, Kirshblum, S, Hearn, J, Razvi, IS, Sikka, S, Callender, L, Bennett, M, Robertson, K, Driver, S, Kline-Quiroz, C, Donovan, J, Botticello, A, Arnold, D, Latham, N, Houlihan, B, Bickmore, T, Trinh, H, Shamekhi, A, Ellis, T, LaVela, SL, Burkhart, E, Kale, I, Bombardier, C, Snoxell, E, Knezevic, S, Hong, EK, Asselin, P, Cirnigliaro, C, Kornfeld, S, Gorman, P, Forrest, G, Bauman, W, Farkas, G, Berg, A, Gater, D, Cleveland, C, Gabet, J, Harrington, A, Arenth, P, Farkas, G, Gorgey, A, Dolbow, D, Berg, A, Gater, D, Luther, S, Finch, D, Bouayad, L, Cirnigliaro, C, Donovan, J, Forrest, G, Gater, D, Kopp, M, Watzlawick, R, Martus, P, Failli, V, Finkenstaedt, F, Chen, Y, DeVivo, M, Dirnagl, U, Schwab, J, LiMonta, J, Santiago, T, Wu, YK, Harel, N, Monden, K, Trost, Z, Nguyen, N, Morse, L, Boals, A, Wenzel, L, Silveira, S, Hughes, R, Nosek, M, LeDoux, T, Taylor, H, Diaz, L, Robinson-Whelen, S, Garshick, E, Betz, K, Krause, J, Cao, Y, Li, C, Hon, B, Kirshblum, S, Khong, CM, Dirlikov, B, Shem, K, Charlifue, S, Song, S & Burns, S 2018, 'Presentation abstracts', The journal of spinal cord medicine, vol. 41, no. 5, pp. 599-622.
Gater, DR, Farkas, GJ, Berg, AS & Castillo, C 2019, 'Prevalence of metabolic syndrome in veterans with spinal cord injury', Journal of Spinal Cord Medicine, vol. 42, no. 1, pp. 86-93.
Farkas, GJ, Gorgey, AS, Dolbow, DR, Berg, AS & Gater, DR 2019, 'Sex dimorphism in the distribution of adipose tissue and its influence on proinflammatory adipokines and cardiometabolic profiles in motor complete spinal cord injury', Journal of Spinal Cord Medicine, vol. 42, no. 4, pp. 430-436.
Liao, JG & Berg, A 2019, 'Sharpening Jensen's Inequality', American Statistician, vol. 73, no. 3, pp. 278-281.
Hammett, E, Veldheer, S, Hrabovsky, S, Yingst, J, Berg, A, Poole, E, Stauffer, D & Foulds, J 2018, 'TXT2STAYQUIT: Pilot randomized trial of brief automated smoking cessation texting intervention for inpatient smokers discharged from the hospital', Journal of Hospital Medicine, vol. 13, no. 7, pp. 488-489.
Chen, P, Hallock, KK, Mulvey, CL, Berg, AS & Cherian, VT 2018, 'The effect of elevated A1C on immediate postoperative complications: A prospective observational study', Clinical Diabetes, vol. 36, no. 2, pp. 128-132.
Farkas, GJ, Gorgey, AS, Dolbow, DR, Berg, AS & Gater, DR 2018, 'The influence of level of spinal cord injury on adipose tissue and its relationship to inflammatory adipokines and cardiometabolic profiles', Journal of Spinal Cord Medicine, vol. 41, no. 4, pp. 407-415.


Crowell, KT, Julian, KG, Katzman, M, Berg, AS, Tinsley, A, Williams, ED, Koltun, WA & Messaris, E 2017, 'Compliance with Clostridium difficile treatment guidelines: Effect on patient outcomes', Epidemiology and Infection, vol. 145, no. 11, pp. 2185-2192.
Wrafter, PF, Connelly, TM, Khan, JSA, Lucey, BC, Berg, A, Koltun, W & Joyce, WP 2017, 'Diverticular disease is associated with benign intra-abdominal cystic disease', Expert Review of Gastroenterology and Hepatology, vol. 11, no. 5, pp. 487-490.
Connelly, TM, Choi, CS, Berg, AS, Harris, L, Coble, J & Koltun, WA 2017, 'Diverticulitis and Crohn's disease have distinct but overlapping tumor necrosis superfamily 15 haplotypes', Journal of Surgical Research, vol. 214, pp. 262-269.
Andrew, RE, Lauria, A, Puleo, FJ, Berg, A & Stewart, DB 2017, 'Inpatient infliximab is ineffective at preventing colectomy for steroid refractory extensive colitis', Journal of Surgical Research, vol. 219, pp. 18-24.
Veldheer, S, Hrabovsky, S, Yingst, J, Sciamanna, C, Berg, A & Foulds, J 2018, 'The Use of Self-Directed Relapse Prevention Booklets to Assist in Maintaining Abstinence After a 6-Week Group Smoking Cessation Treatment Program: A Randomized Controlled Trial', Health Education and Behavior, vol. 45, no. 2, pp. 190-197.

Clinical Trials Search

Children (age < 18 years)
Adults (age >= 18 years)